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Extrinsic factors, such as growth factors and neurotransmitters, secreted by the stem cell niche, diffusing from the nearby vasculature or released from synaptic connections relay the environmental influence onto the progression of neurogenesis in the adult hippocampus (see Figure 1)

By combining the rml1 mutation with mutations in the two genes encoding dual-addressed cytosolic/mitochondrial NADPH-thioredoxin reductases ( NTRA , At2g17420